Condamine Earless Dragon (Tympanocryptis condaminensis)
The Condamine Earless Dragon is a small to medium-sized earless dragon from the Darling Downs and Condamine River region of Queensland. It uses remnant grassland with deep, complex black-soil cracks, including habitat in road reserves, so clearing, earthworks, weed spread and road projects require careful habitat checks.
Conservation status
Commonwealth, EPBC Act 1999: Endangered.
Queensland, Nature Conservation Act 1992: Endangered.
Breeding season
The Condamine Earless Dragon has no documented species-specific breeding calendar. Related grassland earless dragons breed and produce young mainly from spring to autumn, while clearing in winter may harm animals sheltering in soil refuges. No month should be treated as low risk without local survey information.
Reproductive mode: Not recorded for this species. Related grassland earless dragons are described as egg laying.
Clutch size: Not recorded for this species. Related grassland earless dragons are reported to lay 3 to 6 eggs, or up to five eggs per clutch.
Incubation: Not recorded for this species. Related grassland earless dragons have an incubation period of 40 to 70 days.
Size at hatching: Not recorded for this species. Related grassland earless dragon hatchlings may weigh as little as 0.6 grams and measure about 3 cm.
Age at maturity: Not recorded for this species. Related research suggests some grassland earless dragons may mature within their first year, but this is not confirmed.
Nest or egg sites: Not recorded for this species. Related grassland earless dragons lay eggs in invertebrate burrows or shallow scrapes, which are covered with loose soil or small debris.
Breeding frequency: Not recorded for this species. Related research suggests one recruitment period each year, with possible mating in spring and autumn.
Confirm timing locally before clearing, because rainfall, temperature and site conditions may shift breeding, emergence and surface activity, and arrange targeted surveys and qualified herpetological advice where suitable habitat will be disturbed.
- Mating: No mating observations are recorded for the Condamine Earless Dragon. Related grassland earless dragons, including Tympanocryptis lineata and T. osbornei, may mate in autumn and spring.
- Gravid females: No records are available for this species. Related grassland earless dragons have been recorded with gravid females from spring to mid-summer, and from September to January.
- Egg laying: Egg laying has not been recorded for this species. Related grassland earless dragons lay eggs during the spring to summer breeding period, in invertebrate burrows or shallow scrapes.
- Incubation: Incubation has not been measured for this species. Related grassland earless dragons have an incubation period of 40 to 70 days, so eggs laid from September to January may remain in soil or refuges from September to April.
- Hatchlings: Hatchling emergence is not recorded for this species. Related grassland earless dragons hatch or emerge from mid-summer through autumn, with field detections reported from January to May.
- Active period: No species-specific active-season dataset is available, although the species is reported to be diurnal. As a general planning pattern for temperate grassland reptiles, expect most surface activity from spring through autumn, with lower activity in winter.
- Winter inactivity: Brumation or winter inactivity has not been documented for this species. For general planning in temperate grassland habitat, treat May to August as a period when animals may shelter in deep, complex black-soil cracks and be unable to escape machinery.
- Highest clearing risk: Clearing risk is year round because the species may be associated with soil refuges in every season. Risk is especially difficult to avoid from September to May because breeding, eggs and hatchlings may overlap, while June to August may expose animals sheltering in cracks to machinery.
Identification
This is a small to medium-sized, cryptic dragon with strong brown-black and white patterning. It has no visible tympanum, or external ear opening, which is characteristic of earless dragons.
The body has irregular brown-black and white banding and speckling. A narrow, continuous white stripe runs from the axilla to the groin, with a separate narrow white stripe along the back edge of the thigh that extends onto the tail base.
Three pale spots are present on the top of the snout: one above each nostril and one at the snout tip. The limbs are slender and moderately long, and the tail is long, slender and finely tapered.
The head, throat and upper chest commonly have dense dark speckling. Individuals with heavy ventral patterning may have a narrow white stripe along the centre of the upper chest.
Body size: Small to medium-sized, but measurements have not been documented.
Tail: Long and slender, tapering to a fine tip.
External ear: No visible tympanum.
Identification: Diagnostic characters
The dorsal scales are heterogeneous, with unkeeled, weakly keeled and strongly keeled scales interspersed. Scattered enlarged, keeled, pointed scales occur on the head and body.
The species has two preanal pores and no femoral pores. The snout has more than 10 internasals in the diagnosis, with 11 recorded in the holotype.
Preanal pores: Two.
Femoral pores: Absent.
Holotype meristics: 11 supralabials, 10 infralabials, 11 internasals and 10 subocular scales.
Identification: Similar species and field signs
Identification should be checked against other Tympanocryptis species in the region. The contrasting white and brown-black pattern, continuous white lateral stripe, three pale snout spots and absence of a visible tympanum are useful characters, but expert confirmation may be needed.
Species-specific information on tracks, scats, calls, nests, sloughs or eyeshine is limited. The species is cryptic, so absence of direct observations does not demonstrate absence from suitable grassland or road-reserve habitat.
- Photograph the animal from above and below where safe to do so.
- Record the exact location, habitat condition, soil cracks, nearby weeds and whether the animal was found in a road reserve, paddock remnant or other grassland patch.
- Use an experienced reptile ecologist for confirmation where the identification is uncertain.
Distribution: Known range
Confirmed records are from south-east Queensland, centred on the Darling Downs around Oakey and the Bongeen area. The full range remains uncertain, particularly along the Condamine River floodplain south of Chinchilla.
The type series was collected from Oakey, Mt Tyson, Kunari and Brookstead in Queensland. These localities occur in the agricultural Darling Downs region.
There are small areas of native grassland along the Condamine River floodplain south of Chinchilla, with anecdotal reports of earless dragons but no confirmed sightings in the cited study.
State occurrence: Queensland.
Known localities: Oakey, Mt Tyson, Kunari and Brookstead.
Potential range requiring survey: Native grasslands along the Condamine River floodplain south of Chinchilla.
Distribution: Subpopulations and trend
There is no national population estimate, subpopulation count, home-range measurement or quantified population trend.
The species is treated as a significant conservation concern, and further grassland surveys and population genetic work were identified as priorities. Genetic work could help assess connectivity, inbreeding and recent bottlenecks.
Population size: Not provided.
Population trend: Not well quantified.
Connectivity: Requires further population genetic investigation.
Habitat: Grassland habitat
The Condamine Earless Dragon is associated with remnant grassland and specific cracking-soil conditions. Habitat can persist in highly modified agricultural areas and road reserves, but its distribution across remaining grassland is not fully known.
The documented habitat includes remnant grassland patches in the agricultural Darling Downs and possible native grassland along the Condamine River floodplain.
Road reserves can contain suitable habitat. The key structural feature identified for this species is deep, complex black-soil cracking that provides refuge sites.
Habitat type: Remnant native grassland.
Soil feature: Deep, complex black-soil cracks.
Modified settings: Road reserves and other fragmented grassland remnants may support habitat.
Habitat: Habitat condition
Suitable habitat may occur as small, isolated patches within cropland and beside roads. Elevation, rainfall, temperature, vegetation-association and minimum patch-size limits have not been established.
Weed invasion can alter soil structure and reduce the availability of refuge sites. Woody weeds and invasive pasture grasses can shade out the ground features used by these dragons.
- Map native grassland remnants and areas of black cracking soil before clearing or earthworks.
- Inspect road reserves, stock routes, paddock remnants and floodplain grasslands, not only larger intact vegetation patches.
- Record the depth, continuity and condition of soil cracks, together with ground cover and weed cover.
Foraging habitat: Foraging setting
The diet, prey species, feeding times, foraging distances and feeding rates of the Condamine Earless Dragon are not documented. Habitat assessment should therefore retain the grassland structure and soil refuges used by the species while these details remain unresolved.
The species occurs in remnant grassland patches, including fragmented sites in agricultural areas and roadside verges. These areas should be treated as potential foraging habitat where the species or suitable cracking-soil habitat is present.
No specific food plants, prey taxa, foraging range or feeding method were provided.
Diet: Not documented.
Foraging range: Not provided.
Feeding period: Not provided.
- Avoid stripping ground cover and cracking-soil surfaces from remnant grassland patches.
- Limit vehicle movement and soil disturbance within and around known or potential cracking-soil habitat.
- Control weeds using methods that do not destroy soil cracks or introduce further soil disturbance.
Breeding habitat: Breeding biology
Breeding observations and measurements are lacking for this species. Breeding habitat should be assessed within remnant grassland that contains the species' identified soil-refuge features.
Mating, egg laying, clutch size, incubation, age at maturity, lifespan and nest locations are not documented. Avoid assuming that disturbed or apparently vacant grassland is not breeding habitat.
The species' habitat requirements are poorly understood. Site-specific surveys and advice from a qualified herpetologist are appropriate where clearing may affect potential habitat.
Breeding season: Not provided.
Clutch size: Not provided.
Incubation period: Not provided.
Age at maturity: Not provided.
Lifespan: Not provided.
Breeding habitat: Breeding habitat features
Retain remnant native grassland and deep, complex black-soil cracks while breeding use is being assessed. Nest-site dimensions, preferred vegetation height, bare-ground cover and other breeding-site measurements are not well documented.
- Schedule targeted reptile surveys before clearing in suitable grassland and cracking-soil habitat.
- Protect habitat patches and soil cracks from grading, trenching, compaction and weed-driven structural change.
- Treat connected remnants and road-reserve grasslands as potential breeding habitat until survey information shows otherwise.
Sheltering habitat: Soil refuges
Deep, complex cracks in black soils are the only species-specific refuge feature identified. These cracks can occur in remnant grassland and road reserves and may be damaged by soil disturbance or weeds.
The species occupies remnant grassland patches containing deep, complex black-soil cracks. These cracks are identified as refuge sites and may be important in small grassland remnants and road reserves.
Crack depth, width, density and distance from vegetation cover have not been measured.
Known refuge: Deep, complex cracks in black soil.
Refuge dimensions: Not provided.
Other shelters: No species-specific information was provided on burrows, hollows, logs, rocks or litter.
- Keep heavy machinery, spoil, trenching and temporary access tracks away from cracking-soil habitat.
- Inspect cracks before disturbance and record their distribution, condition and connectivity across the site.
- Retain native ground cover around cracks and prevent soil compaction.
Sheltering habitat: Weed-related refuge loss
Weed invasion can change soil structure and reduce refuge availability. Lantana and invasive pasture grasses such as buffel grass can shade out critical refuge sites.
- Clean machinery and vehicles before entering suitable grassland to reduce weed spread.
- Control weeds before earthworks where practicable, using methods that protect cracking soils.
- Do not stockpile soil, mulch or vegetation over known or potential refuge cracks.
Threats: Clearing and fragmentation
The main documented risks are habitat loss and fragmentation, degradation of cracking-soil refuges, weed invasion and infrastructure disturbance. The species' Endangered status means small remnant patches should be assessed before clearing.
Clearing remnant grassland can remove habitat and divide already fragmented populations. The species may occur in small patches, croplands and roadside verges, so apparently narrow or isolated remnants should not be dismissed.
The full distribution is uncertain. Unconfirmed reports south of Chinchilla show that additional grassland surveys are needed before habitat is cleared.
- Survey native grassland, road reserves and black cracking soils before clearing.
- Avoid clearing known habitat and retain links between remnant patches where possible.
- Use a qualified ecologist to assess potential habitat when no animal is detected.
Threats: Weeds and soil disturbance
Transport construction can spread weeds through earthworks, soil transport, poor weed hygiene and vehicle movement. Weed invasion can alter soil structure and reduce deep crack refuges.
Lantana and buffel grass can shade critical refuge sites. Grading, excavation, compaction and spoil placement can also directly damage cracking soils.
- Apply weed hygiene controls to all vehicles, machinery, soil and imported materials.
- Keep construction compounds, access tracks and spoil outside cracking-soil grassland wherever practicable.
- Inspect and maintain exclusion fencing around retained habitat.
Threats: Roads and construction
Road projects can remove roadside habitat and create barriers between remnants. Construction noise and vibration may affect reptiles, although the extent of these effects for Queensland reptiles is not known.
The transport guidance recommends avoiding critical reptile habitat where possible. Land bridges can provide continuous habitat and shelter, while no reptile-specific underpasses have been installed in Australia according to the cited guidance.
- Prioritise habitat avoidance over later mitigation.
- Use specialist advice for crossing structures or barrier treatments where a project cannot avoid habitat.
- Manage vehicle access and construction timing around confirmed animals and retained habitat.
Threats: Fire, predators, disease and hydrology
Species-specific evidence on fire, predators, disease or hydrology change is limited. Assess these pressures at the site level, particularly where fire, floodplain works or changed drainage may alter grassland structure and black-soil cracking.
Do not infer that these pressures are absent because evidence of them may be incomplete.
- Record recent fire history, planned burns, drainage changes, floodplain works and predator activity during site assessment.
- Seek species-specific advice before using fire or altering surface water movement in potential habitat.
Survey methodology: 1. Habitat assessment and systematic searches
No species-specific survey protocol, detection probability, seasonal window or minimum effort has been established for the Condamine Earless Dragon. Surveys should therefore be designed by a suitably experienced herpetologist, with effort matched to the area and condition of remnant grassland and cracking soil habitat.
Map remnant grassland patches, roadside verges and other areas with deep, complex black soil cracks before selecting survey methods.
Inspect cracking soils, ground cover and refuge sites systematically because the species is associated with remnant grassland patches containing deep, complex black soil cracks.
Search the full proposed disturbance area and connected retained habitat where access allows, and record all observations, likely refuge sites and barriers.
Survey methodology: 2. Active reptile searches
Use experienced observers to search suitable ground refuges and grassland structure during conditions selected to maximise reptile activity and detection.
Search around soil cracks and other ground refuges without collapsing or filling the cracks before the fauna management plan is approved.
Record the date, start and finish time, weather, ground conditions, search area, observer effort, detection method and exact location for every animal or sign detected.
Survey methodology: 3. Night searches and trapping
Consider night searches and approved trapping methods where daytime searches are unlikely to detect the species, because the dragon is cryptic and no single reliable method has been established.
Set the number of nights, search hours, transect length, trap nights and trap spacing in a project-specific survey plan prepared by a qualified herpetologist.
Do not use trapping or handling unless the required animal ethics, state wildlife and project approvals are in place, and ensure traps are checked and animals are released promptly under the approved method.
Survey methodology: Minimum effort and reporting
- Use a documented, repeatable survey design that covers all suitable habitat and explains why the selected methods and effort are adequate, because no species-specific minimum effort has been established.
- Record negative as well as positive results, including the area searched, habitat condition, weather and observer effort.
- Submit confirmed records, with coordinates and supporting evidence, to the relevant Queensland wildlife database and any project or regulatory database required by the approval.
- Review the survey design with a species expert if habitat is fragmented, access is restricted or detection is expected to be low.
Vegetation associations: Tussock grassland
Source-checked associations and automatically indexed candidates are separated below. An association does not establish site presence, preference or dependence.
The documented habitat includes remnant grassland patches in the agricultural Darling Downs and possible native grassland along the Condamine River floodplain.
Automatically indexed from existing profile text. Not a verified preference, mapped occurrence or habitat requirement. Check the original source and regional qualifiers before use.
Evidence: Unreviewed profile-derived candidate
Confidence: Not independently verified
Region: Scope as described in the quoted profile text; not independently verified. Profile states: QLD.
Habitat use: general
Attribution gap: No sentence-level reference is attached to the original profile block. Review the profile bibliography; this candidate is not source-verified.
Ecosystem associations
Source-checked associations and automatically indexed candidates are separated below. An association does not establish site presence, preference or dependence.
No species-specific ecosystem association has been indexed or source-checked in this evidence library. This is an evidence gap, not evidence of no association.
Soil and substrate associations: Cracking clays or gilgai
Source-checked associations and automatically indexed candidates are separated below. An association does not establish site presence, preference or dependence.
It uses remnant grassland with deep, complex black-soil cracks, including habitat in road reserves, so clearing, earthworks, weed spread and road projects require careful habitat checks.
Automatically indexed from existing profile text. Not a verified preference, mapped occurrence or habitat requirement. Check the original source and regional qualifiers before use.
Evidence: Unreviewed profile-derived candidate
Confidence: Not independently verified
Region: Scope as described in the quoted profile text; not independently verified. Profile states: QLD.
Habitat use: general
Attribution gap: No sentence-level reference is attached to the original profile block. Review the profile bibliography; this candidate is not source-verified.
Water and wetland associations: Water-associated habitat mentioned in profile
Source-checked associations and automatically indexed candidates are separated below. An association does not establish site presence, preference or dependence.
The documented habitat includes remnant grassland patches in the agricultural Darling Downs and possible native grassland along the Condamine River floodplain.
Automatically indexed from existing profile text. Not a verified preference, mapped occurrence or habitat requirement. Check the original source and regional qualifiers before use.
Evidence: Unreviewed profile-derived candidate
Confidence: Not independently verified
Region: Scope as described in the quoted profile text; not independently verified. Profile states: QLD.
Habitat use: general
Attribution gap: No sentence-level reference is attached to the original profile block. Review the profile bibliography; this candidate is not source-verified.
Fauna records and mapped habitat: Not yet analysed
Record locations can be overlaid with mapping, but record frequency alone does not measure habitat preference. No species is excluded from desktop assessments by this analysis.
No quality-controlled record-to-map analysis has been completed for this species. The existing generalised distribution dots and indicative range polygons have not been treated as precise occurrence coordinates.
A defensible run needs dated, independently attributable records, coordinate uncertainty, duplicate and captive-record checks, layer versions and regional scope. Preference and absence require additional survey-effort and detection evidence.
Habitat limitations and contrary evidence: No supported exclusion recorded
Not recorded is not the same as cannot occur. Low suitability, resident habitat, breeding habitat and occasional movement are different conclusions.
This library has not established a vegetation, ecosystem or soil-type exclusion for this species. This does not mean all habitat is suitable; the evidence needed to justify an exclusion has not been assembled.
Habitat limitations and contrary evidence: Before using a negative association
- Confirm the taxon, geographic region, life stage and habitat function covered by the evidence.
- Check survey methods, season, repeated effort and detectability before interpreting non-detection.
- Investigate credible contradictory records and microhabitats absent from broad mapping.
- Do not convert a missing map class, low record count or failed data request into species absence.
Before habitat disturbance: Avoid and minimise
Avoiding the species' habitat is the preferred control for transport and construction works. Focus planning on remnant grassland, deep cracking black soils, refuge sites and connectivity between retained patches.
Redesign the footprint to retain remnant grassland patches with deep, complex black soil cracks, including suitable habitat in road reserves.
Retain links between habitat remnants where practicable, because the species may occur in highly fragmented populations and research indicates a need to understand habitat connectivity.
Keep machinery, spoil, access tracks and laydown areas out of cracking soil habitat and roadside refuge areas.
Prevent changes to soil structure and avoid shading or covering cracks with fill, stockpiles or invasive vegetation.
Before habitat disturbance: Approvals and work planning
Check whether the proposed action requires referral or approval under the EPBC Act because the species is listed as Endangered nationally.
Obtain the relevant Queensland wildlife permits and any other state or territory approvals before survey, trapping, handling or relocation.
Prepare a fauna management plan that identifies habitat to retain, survey methods, authorised handlers, stop-work triggers, release locations and incident reporting.
Do not rely on a generic reptile procedure where the project affects deep cracking soils or known or suspected Condamine Earless Dragon habitat.
Before habitat disturbance: Site preparation
Complete the approved pre-clearance survey immediately before works and update the habitat map if soil cracks, grassland remnants or refuge sites are found.
Clearly mark no-go areas, retained habitat, access routes and any exclusion zones before machinery enters the site.
Install exclusion fencing where it can prevent entry into retained habitat without damaging cracking soils or blocking animal movement between retained patches.
Control weed movement through clean machinery, clean soil handling and weed hygiene because transport works can spread weeds through earthworks, soil movement and vehicles.
During habitat disturbance: Daily controls and machinery interface
Treat each work front as a controlled fauna operation. The fauna spotter catcher must have authority to stop work when an animal or suitable refuge habitat is found.
Brief all workers each day on the species, mapped habitat, exclusion zones, contact procedure and stop-work authority.
Inspect the work area before machinery starts and after breaks, relocations or changes to the work front.
Keep the fauna spotter catcher in a position where the operator can see and respond to instructions, without placing the spotter in an unsafe position.
Use staged clearing so the spotter catcher can inspect each section before disturbance proceeds.
Prevent vehicles and machinery from spreading weeds or soil into retained cracking-soil habitat.
During habitat disturbance: Habitat and animal handling
Stop work if a Condamine Earless Dragon is found, if a potentially occupied crack or other refuge cannot be safely inspected, or if clearing begins outside the approved footprint.
Allow only authorised and appropriately permitted personnel to capture, handle or relocate the animal.
Move an animal only when necessary to prevent immediate harm, and release it into nearby retained habitat selected in the fauna management plan.
Do not place an animal in isolated habitat or across a road, active work area or other barrier to movement.
Transfer an injured animal to an appropriately experienced veterinarian or authorised wildlife carer as soon as practicable.
During habitat disturbance: Records and incident response
Record every sighting, capture, relocation, injury, mortality, stop-work event and release location with coordinates, time, habitat description and photographs where safe.
Notify the project environmental officer and the relevant approval agency of incidents within the time required by the approval or fauna management plan.
Reassess the work method if repeated animals, occupied refuge sites or unexpected habitat are found.
Keep daily records of cleared area, personnel, machinery, inspections, exclusion fencing and weed hygiene checks.
After habitat disturbance: Post-clearance checks
Post-clearance work should confirm that retained habitat and movement routes remain intact. Key management concerns include habitat loss, fragmentation, soil change and weed invasion.
Inspect the cleared edge and retained cracking-soil habitat for animals, damaged refuge sites, soil movement, shading and weed establishment.
Confirm that no-go areas, exclusion fencing and access controls remain in place while construction continues.
Repair or remove controls only under the fauna management plan and approval conditions.
After habitat disturbance: Monitoring and reporting
Monitor retained habitat and any relocated animals using the approved project method and schedule, because no species-specific monitoring period or success threshold has been established.
Compare post-clearance observations with pre-clearance survey records and investigate any decline in habitat condition or detections.
Submit confirmed records and mortality data to the relevant Queensland wildlife database and report against EPBC, state and project approval conditions.
After habitat disturbance: Offsets and corrective action
Apply any offset, habitat protection or restoration requirement specified by the EPBC Act decision, Queensland approval or other project approval.
Escalate weed invasion, soil compaction, loss of cracks or severed connectivity to the environmental officer and implement corrective action under the approved plan.
Use further expert survey or genetic and connectivity investigation where the project may isolate remnant populations.
Rehabilitation actions: Restore habitat structure
Rehabilitation should restore the ground conditions used by this species rather than relying on tree planting alone. Focus on native grassland, deep complex black soil cracks, refuge availability, weed control and habitat connectivity.
Re-establish native grassland across disturbed areas where the original community can be identified, using locally appropriate species and seed sources selected by the restoration ecologist.
Reinstate natural soil profiles and drainage, and do not seal, fill or compact existing or recreated cracking-soil habitat.
Reconnect restored areas with retained grassland where practicable, while avoiding new barriers from roads, fencing, stockpiles and ongoing vehicle access.
Do not install logs, rocks, hollows or nest boxes as a substitute for the deep cracking-soil refuges identified for this species.
Rehabilitation actions: Weed and disturbance control
Inspect rehabilitation areas for woody weeds and invasive pasture grasses, including Lantana and buffel grass, which can shade refuge sites and alter soil conditions.
Use clean machinery and soil handling procedures during revegetation to prevent weed spread into retained and restored habitat.
Manage grazing, vehicle access and other activities that compact soil or reduce the availability of cracks and ground refuges.
Rehabilitation actions: Fire and monitoring
Set fire management actions with the land manager and local ecologist so that fire does not remove the grassland structure or expose restored cracking soils to erosion.
Monitor native grass cover, weed cover, soil cracking, refuge availability, connectivity and dragon detections against baseline records.
Define measurable completion criteria in the rehabilitation plan before works begin, because species-specific rehabilitation targets and monitoring timeframes have not been published.
Revise weed control, access management or restoration methods if monitoring shows declining habitat condition or continued fragmentation.
Sources
- The Taxonomic Status of Endangered Earless Dragons ..., pmc.ncbi.nlm.nih.gov: https://pmc.ncbi.nlm.nih.gov/articles/PMC4116116/
- Fauna Sensitive Transport Infrastructure Delivery Chapter ..., Queensland Government: https://www.tmr.qld.gov.au/_/media/busind/techstdpubs/environment-management/fauna-sensitive-transport-infrastructure-delivery-manual/fstid-chapter-17---reptiles.pdf
- Tympanocryptis for Kids, kids.kiddle.co: https://kids.kiddle.co/Tympanocryptis
- Population structure, growth and reproduction in two species of Tympanocryptis (Agamidae), publish.csiro.au: https://www.publish.csiro.au/zo/pdf/ZO24034
- Condamine Earless Dragon (Tympanocryptis condaminensis), allanglescreatures.com: https://www.allanglescreatures.com/species/tympanocryptis-condaminensis
- About The Dragons - Grassland Earless Dragon Alliance, earlessdragonalliance.com.au: https://earlessdragonalliance.com.au/about-the-dragons/
- Atlas of Living Australia: scatterplot case study and survey-effort limitations, Habitat evidence reference: https://support.ala.org.au/support/solutions/articles/6000208426-scatterplot-case-study